A New Specis of the Isolated Psaroniaceous Rachis from the 一种新的分离psaroniaceous轴从.docVIP

A New Specis of the Isolated Psaroniaceous Rachis from the 一种新的分离psaroniaceous轴从.doc

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A New Specis of the Isolated Psaroniaceous Rachis from the 一种新的分离psaroniaceous轴从

A New Species of the Isolated Psaroniaceous Rachis from the Early Permian in China Ning Xiang 1,2, Shi-Min Ma 3, Bao-Lin Tian 3 and Shi-Jun Wang 1* State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, the Chinese Academy of Sciences, Beijing 100093, China; 2. Graduate University of the Chinese Academy of Sciences, Beijing 100049, China; 3. China University of Mining and Technology, Beijing 100083, China) Received : 12 Oct.2006 Accepted 24 Apr. 2007 Handling editor: Cheng-Sen Li Category: Evolution Author for correspondence . Tel:+86 (0) 1062836435 E-mail:wangsj@ibcas.ac.cn Supported by the National Natural Foundation of China and the Special Fund from the Director of Institute of Botany, CAS. Abstract A new kind of marattialean raches are reported from the coal balls in Coal Seam No.7 in the upper part of Taiyuan Formation (early Early Permian) from Taiyuan, Shanxi, China and are assigned to the genus Stipitopteris Grand’Eury (Psaroniaceae). The present specimens are different from all the six reported species of the genus thus to be erected as a new species: Stipitopteris shanxiensis. The raches of the new species are generally dorsi-ventrally flattened. Main raches usually exhibit scales of different forms on their surface. Beneath the epidermis is a zone of parenchymatous cells some of which contain tannin-like contents. Inside this is a zone of small sclerenchymatous cells. Inward are the ground tissue and vascular bundles. The vascular bundles are continuous and are in two circles: outer circle assumes a transversely elliptical shape with the gap and pinna trace; and inner circle assumes a shallow “C” shape with enrolled ends. The ground tissue located at the inner side of the vascular bundle is composed of thicker-walled parenchymatous cells. The cells of the ground tissue are vertically elongated in longitudinal section. Subordered raches are smaller and have simpler structures than the main raches. T

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